African Dung Beetles Online

520 species of dung beetles (Scarabée bousier) are known to occur in Democratic Republic of Congo XXX of which are endemic. If you are aware of other species not included here please e-mail christopherborrow@gmail.com.

• ALLOGYMNOPLEURUS
  
  • Allogymnopleurus alluaudi Garreta, 1914 (BMNH drawer 34.1)
  • Allogymnopleurus chloris Klug, 1855 (BMNH drawer 34)
  • Allogymnopleurus umbrinus Gerstaecker, 1871 (BMNH drawer 34.1)
• ALLONITIS
  
  • Allonitis nasutus Felsche, 1907 (BMNH drawer 172)
• ALLOSCELUS
  
  • Alloscelus paradoxus Boucomont, 1923 (BMNH drawer 43)
• ANACHALCOS
  
  • Anachalcos aurescens Bates, 1888 (BMNH drawer 64)
  • Anachalcos convexus Boheman, 1857 (BMNH drawer 63)
  • Anachalcos cupreus Fabricius, 1775 (BMNH drawer 63)
  • Anachalcos procerus Gerstaecker, 1874 (BMNH drawer 64)
• CACCOBIOMORPHUS
  
  • Caccobiomorphus congolanus Balthasar, 1964
• CACCOBIUS
  
  • Caccobius pentagonus d'Orbigny, 1908 (BMNH drawer 202)
• CACCOPHILUS
  
  • Caccophilus convexifrons Raffray, 1877 (BMNH drawer 203)
  • Caccophilus cribrarius Boucomont, 1928 (BMNH drawer 202)
  • Caccophilus discrepans Peringuey, 1901 (BMNH drawer 201)
  • Caccophilus ferrugineus Fahraeus, 1857 (BMNH drawer 201)
  • Caccophilus gibbosulus d'Orbigny, 1915
  • Caccophilus globaticeps d'Orbigny, 1905 (BMNH drawer 202)
  • Caccophilus grossegranosus Balthasar, 1963
  • Caccophilus hirsutus Frey, 1958
  • Caccophilus inops Peringuey, 1901
  • Caccophilus leleupi Balthasar, 1963
  • Caccophilus obtusus Fahraeus, 1857 (BMNH drawer 201)
  • Caccophilus ocellipennis d'Orbigny, 1913 (BMNH drawer 201)
  • Caccophilus pilosus Frey, 1958 (BMNH drawer 264)
  • Caccophilus postulatus d'Orbigny, 1905 (BMNH drawer 203)
  • Caccophilus punctatissimus Harold, 1867 (BMNH drawer 201)
  • Caccophilus reticuliger d'Orbigny, 1904 (BMNH drawer 201)
  • Caccophilus schaedlei d'Orbigny, 1902 (BMNH drawer 203)
  • Caccophilus seminulum Klug, 1855 (BMNH drawer 203)
  • Caccophilus upembanus Frey, 1958
• CATHARSIUS
  
  • Catharsius achates Olivier, 1789 (BMNH drawer 115)
  • Catharsius alpheus Gillet, 1933 (BMNH drawer 116)
  • Catharsius approximans Kolbe, 1893
  • Catharsius balubanus Kolbe, 1893
  • Catharsius biconifer Felsche, 1907 (BMNH drawer 117)
  • Catharsius bicornutus Kolbe, 1893
  • Catharsius brevicornis Felsche, 1907 (BMNH drawer 116)
  • Catharsius brittoni Ferreira, 1963 (BMNH drawer 116)
  • Catharsius cassius Kolbe, 1893
  • Catharsius congolensis Ferreira, 1962
  • Catharsius duciformis Ferreira, 1959
  • Catharsius dux Harold, 1878 (BMNH drawer 113)
  • Catharsius garambae Ferreira, 1962
  • Catharsius gorilla Thomson, 1858 (BMNH drawer 113)
  • Catharsius gorilloides Felsche, 1907
  • Catharsius kolbei Felsche, 1907
  • Catharsius luluensis Ferreira, 1960
  • Catharsius machadoi Ferreira, 1960
  • Catharsius neptunus Kolbe, 1893 (BMNH drawer 117)
  • Catharsius ninus Gillet, 1918
  • Catharsius pandion Harold, 1877
  • Catharsius platicornis Ferreira, 1960
  • Catharsius platycerus Klug, 1855 (BMNH drawer 115)
  • Catharsius platynotus Felsche, 1907 (BMNH drawer 115)
  • Catharsius polynices Kolbe, 1893 (BMNH drawer 115)
  • Catharsius pseudooedipus Ferreira, 1960
  • Catharsius saegeri Ferreira, 1960
  • Catharsius satyrus Kolbe, 1893 (BMNH drawer 114)
  • Catharsius sesostris Waterhouse, 1888 (BMNH drawer 118)
  • Catharsius spectabilis Gillet, 1918 (BMNH drawer 113)
  • Catharsius straeleni Ferreira, 1963
  • Catharsius stuhlmanni Kolbe, 1893 (BMNH drawer 116)
  • Catharsius tricornutus Degeer, 1778 (BMNH drawer 117)
  • Catharsius upembanus Ferreira, 1962
  • Catharsius vitulus Boheman, 1857 (BMNH drawer 117)
• COPRIDASPIDUS
  
  • Copridaspidus peregrinus Harold, 1878 (BMNH drawer 142)
• COPRIS
  
  • Copris amabile Ferreira, 1959
  • Copris arcturus Gillet, 1907 (BMNH drawer 132)
  • Copris armiger Gillet, 1910 (BMNH drawer 137)
  • Copris atropolitus Gillet, 1910 (BMNH drawer 133)
  • Copris basilewskyi Ferreira, 1962
  • Copris bovinus Gillet, 1908 (BMNH drawer 137)
  • Copris carmelita Fabricius, 1801 (BMNH drawer 134)
  • Copris colmanti Gillet, 1908 (BMNH drawer 134)
  • Copris dracunculus Fereira, 1959
  • Copris elphenor Klug, 1855 (BMNH drawer 131)
  • Copris fidius Olivier 1789 (BMNH drawer 133)
  • Copris gazellarum Gillet 1918 (BMNH drawer 133)
  • Copris inhalatus Quedenfeldt 1884 (BMNH drawer 132)
  • Copris insidiosus Peringuey 1901 (BMNH drawer 137)
  • Copris integer Reische 1847 (BMNH drawer 132)
  • Copris interioris Kolbe 1897 (BMNH drawer 134)
  • Copris jucundus Gillet 1933
  • Copris laioides Boucomont 1932 (BMNH drawer 131)
  • Copris laius Harold 1868 (BMNH drawer 131)
  • Copris megaceratoides Waterhouse 1891 (BMNH drawer 132)
  • Copris mesacanthus Harold 1878 (BMNH drawer 133)
  • Copris misellus Peringuey 1901 (BMNH drawer 136)
  • Copris moffartsi Gillet 1907 (BMNH drawer 133)
  • Copris nepos Gillet 1908 (BMNH drawer 136)
  • Copris orion Klug 1835 (BMNH drawer 136)
  • Copris orphanus Guerin 1847 (BMNH drawer 137)
  • Copris overlaeti Balthasara 1961
  • Copris phylax Gillet 1908
  • Copris tridens Felsche 1901 (BMNH drawer 134)
  • Copris wiesei Kolbe 1914 (BMNH drawer 137)
• COPTORHINA
  
  • Coptorhina auspicata Peringuey 1901 (BMNH drawer 71)
  • Coptorhina davidi Janssens 1939
  • Coptorhina nitidipennis Boheman 1857 (BMNH drawer 71)
  • Coptorhina optatus Peringuey 1901 (BMNH drawer 70)
  • Coptorhina subaenea Janssens 1939
• CYPTOCHIRUS
  
  • Cyptochirus distinctus Janssens 1953
• DELOPLEURUS
  
  • Delopleurus giletti Janssens 1939
• DIASTELLOPALPUS
  
  • Diastellopalpus balubanus Kolbe 1893
  • Diastellopalpus basilobatus d'Orbigny 1902 (BMNH drawer 210)
  • Diastellopalpus conradti d'Orbigny 1902
  • Diastellopalpus gilleti d'Orbigny 1908 (BMNH drawer 209)
  • Diastellopalpus infernalis Lansberge 1887 (BMNH drawer 209)
  • Diastellopalpus johnstoni Waterhouse 1885 (BMNH drawer 210)
  • Diastellopalpus lamellicollis Quedenfeldt 1884 (BMNH drawer 209)
  • Diastellopalpus murrayi Harold 1868 (BMNH drawer 210)
  • Diastellopalpus noctis Thomson 1858 (BMNH drawer 209)
  • Diastellopalpus sulciger Kolbe 1893 (BMNH drawer 210)
  • Diastellopalpus tridens Fabricius 1781 (BMNH drawer 209)
• DIGITONTHOPHAGUS
  
  • Digitonthophagus gazella Fabricius 1787 (BMNH drawer 221)
• DREPANOCERUS
  
  • Drepanocerus abyssinicus Roth 1851 (BMNH drawer 186)
  • Drepanocerus bechynei Janssens 1953
  • Drepanocerus caelatus Gerstaecker 1871 (BMNH drawer 186)
  • Drepanocerus fastiditus Peringuey 1901 (BMNH drawer 186)
  • Drepanocerus kirbyi Kirby 1828 (BMNH drawer 186)
  • Drepanocerus laticollis Fahraeus 1857 (BMNH drawer 186)
  • Drepanocerus marshalli Boucomont 1921 (BMNH drawer 186)
  • Drepanocerus parallelus Raffray 1877 (BMNH drawer 186)
  • Drepanocerus saegeri Balthasar 1963
  • Drepanocerus strigatus Janssens 1953
  • Drepanocerus sulcicollis Castelnau 1840 (BMNH drawer 186)
• EUONITICELLUS
  
  • Euoniticellus fumigatus Boucomont 1923
  • Euoniticellus intermedius Reiche 1849 (BMNH drawer 188)
  • Euoniticellus kawanus Janssens 1939
  • Euoniticellus nasicornis Reiche 1849 (BMNH drawer 190)
  • Euoniticellus parvus Kraatz 1895 (BMNH drawer 188)
  • Euoniticellus tibatensis Kolbe 1904 (BMNH drawer 188)
  • Euoniticellus triangulatus Harold 1873 (BMNH drawer 190)
  • Euoniticellus wittei Janssens 1938
• EUONTHOPHAGUS
  
  • Euonthophagus carbonarius Klug 1855 (BMNH drawer 220)
• EUSAPROECIUS
  
  • Eusaproecius congolanus Balthasar 1963
  • Eusaproecius magnigena d'Orbigny 1913 (BMNH drawer 70)
  • Eusaproecius punctulatus Boucomont 1928
  • Eusaproecius tinantae Boucomont 1923 (BMNH drawer 70)
• FURCONTHOPHAGUS
  
  • Furconthophagus variegatus Fabricius 1798 (BMNH drawer 231)
• GARRETA
  
  • Garreta basilewskyi Balthasar 1961
  • Garreta crenulatus Kolbe 1895 (BMNH drawer 32)
  • Garreta malleolus Kolbe 1895 (BMNH drawer 33)
  • Garreta nitens Olivier 1789 (BMNH drawer 33.1)
• GYMNOPLEURUS
  
  • Gymnopleurus colmanti Janssens 1938 (BMNH drawer 38.1)
  • Gymnopleurus fulgidus Olivier 1789 (BMNH drawer 36)
  • Gymnopleurus jacksoni Waterhouse 1890 (BMNH drawer 38.1)
  • Gymnopleurus sericeifrons Fairmaire 1887 (BMNH drawer 37)
  • Gymnopleurus virens Erichson 1843 (BMNH drawer 37)
• HAROLDIUS
  
  • Haroldius ennearthrus Janssens 1949
  • Haroldius leleupi Janssens 1953
  • Haroldius modestus Janssens 1953
• HELIOCOPRIS
  
  • Heliocopris andersoni Bates 1868
  • Heliocopris anguliceps Janssens 1943
  • Heliocopris antenor Olivier 1789 (BMNH drawer 86)
  • Heliocopris colossus Bates 1868 (BMNH drawer 79)
  • Heliocopris corniculatus Janssens 1940
  • Heliocopris dolosus Janssens 1940
  • Heliocopris erycoides Felsche 1907 (BMNH drawer 85)
  • Heliocopris eryx Fabricius 1801 (BMNH drawer 85)
  • Heliocopris felschei Kolbe 1904
  • Heliocopris haroldi Kolbe 1893 (BMNH drawer 84)
  • Heliocopris helleri Felsche 1907 (BMNH drawer 85)
  • Heliocopris japetus Klug 1855 (BMNH drawer 80)
  • Heliocopris kolbei Felsche 1901 (BMNH drawer 84)
  • Heliocopris mimus Janssens 1940
  • Heliocopris minos Gillet 1907 (BMNH drawer 84)
  • Heliocopris mutabilis Kolbe 1893
  • Heliocopris neptunoides Janssens 1940
  • Heliocopris pauliani Janssens 1940
  • Heliocopris quinqueangulatus Janssens 1940
  • Heliocopris samson Harold 1878
  • Heliocopris staudingeri Kolbe 1893 (BMNH drawer 86)
  • Heliocopris sylvanus Gillet 1925
• HETEROCLITOPUS
  
  • Heteroclitopus burgeoni Boucomont 1926
  • Heteroclitopus collarti Janssens 1939
• HETERONITIS
  
  • Heteronitis castelnaui Harold 1862 (BMNH drawer 174)
• HOWDEN & GILLETELLUS
  
  • Howden & Gilletellus collarti Janssens 1937
• HYALONTHOPHAGUS
  
  • Hyalonthophagus pseudoalcyon d'Orbigny 1913 (BMNH drawer 217)
• KHEPER
  
  • Kheper cupreus Castelnau 1840 (BMNH drawer 17)
  • Kheper festivus Harold 1868 (BMNH drawer 20)
  • Kheper intermedius Gillet 1909 (BMNH drawer 20)
  • Kheper lamarcki Macleay 1821 (BMNH drawer 21)
  • Kheper prodigiosus Erichson 1843 (BMNH drawer 21)
  • Kheper pustulosus Gerstaecker 1871 (BMNH drawer 18)
  • Kheper subaeneus Harold 1869 (BMNH drawer 22)
• LIATONGUS
  
  • Liatongus arrowi Boucomont 1921 (BMNH drawer 194)
  • Liatongus aterrimus Boucomont 1921
  • Liatongus ferreirae Balthasar 1964
  • Liatongus fulvostriatus d'Orbigny 1916 (BMNH drawer 194)
  • Liatongus interruptus Quedenfeldt 1884 (BMNH drawer 193)
  • Liatongus militaris Castelnau 1840 (BMNH drawer 194)
  • Liatongus schoutedeni Boucomont 1920
  • Liatongus sjostedti Felsche 1904 (BMNH drawer 194)
  • Liatongus spathulatus Roth 1851 (BMNH drawer 194)
  • Liatongus taurus Boucomont 1920
  • Liatongus testudo Lansberge 1886 (BMNH drawer 194)
  • Liatongus upembanus Janssens 1953
• LITOCOPRIS
  
  • Litocopris muticus Boheman 1857 (BMNH drawer 137)
  • Litocopris punctiventris Waterhouse 1891 (BMNH drawer 137)
• METACATHARSIUS
  
  • Metacatharsius abortivus Fairmaire 1891 (BMNH drawer 121)
  • Metacatharsius basilewskyi Balthasar 1961
  • Metacatharsius bidentatus Ferreira 1964
  • Metacatharsius kindiai Paulian 1939
  • Metacatharsius modestus Balthasar 1965
  • Metacatharsius peleus Olivier 1789 (BMNH drawer 120)
  • Metacatharsius perpunctatus Balthasar 1965
  • Metacatharsius petrovitzi Balthasar 1965
  • Metacatharsius seminulum Balthasar 1940
  • Metacatharsius simulator Balthasar 1939
  • Metacatharsius tchadensis Balthasar 1961
  • Metacatharsius tricarenatus Ferreira 1964
  • Metacatharsius useramus Kolbe 1893
  • Metacatharsius wittei Ferreira 1964
• MILICHUS
  
  • Milichus apicalis Fahraeus 1857 (BMNH drawer 204)
  • Milichus inaequalis Boucomont 1928
  • Milichus incisiceps Janssens 1954
  • Milichus picticollis Gerstaecker 1871 (BMNH drawer 204)
  • Milichus rhodesianus Peringuey 1904 (BMNH drawer 204)
• MIMONTHOPHAGUS
  
  • Mimonthophagus apicehirtus d'Orbigny 1915 (BMNH drawer 219)
• NEOSAPROECIUS
  
  • Neosaproecius trituberculatus Frey 1958
• NEOSISYPHUS
  
  • Neosisyphus basilewskyi Haff (BMNH drawer 42.4)
  • Neosisyphus fortuitus Peringuey 1901 (BMNH drawer 42.3)
  • Neosisyphus gladiator Arrow 1927 (BMNH drawer 42.3)
  • Neosisyphus rugosus Gory 1833 (BMNH drawer 42.3)
  • Neosisyphus spinipes Thunberg 1818 (BMNH drawer 42.3)
• ONITICELLUS
  
  • Oniticellus egregius Klug 1855 (BMNH drawer 191)
  • Oniticellus pictus Hausmann 1807 (BMNH drawer 191)
  • Oniticellus planatus Castelnau 1840 (BMNH drawer 191)
  • Oniticellus pseudoplanatus Balthasar 1964
• ONITIS
  
  • Onitis adelphus Kolbe 1907 (BMNH drawer 180)
  • Onitis aeneus Lansberge 1875 (BMNH drawer 180)
  • Onitis aerarius Harold 1878 (BMNH drawer 184)
  • Onitis aeruginosus Klug 1855 (BMNH drawer 184)
  • Onitis alexis Klug 1835 (BMNH drawer 181)
  • Onitis androcles Janssens 1937 (BMNH drawer 183)
  • Onitis artuosus Gillet 1909 (BMNH drawer 184)
  • Onitis consaguineus Felsche 1911
  • Onitis cryptodus Gillet 1918 (BMNH drawer 183)
  • Onitis cupreus Castelnau 1840 (BMNH drawer 184)
  • Onitis deceptor Peringuey 1901 (BMNH drawer 179)
  • Onitis fabricii Roth 1851 (BMNH drawer 179)
  • Onitis laminifer Gillet 1909
  • Onitis lycophron Klug 1855 (BMNH drawer 180)
  • Onitis mniszechianus Janssens 1951 (BMNH drawer 184)
  • Onitis monstrosus Felsche 1910 (BMNH drawer 184)
  • Onitis nemoralis Gillet 1932 (BMNH drawer 183)
  • Onitis nubiensis Lansberge 1875 (BMNH drawer 179)
  • Onitis overlaeti Janssens 1938
  • Onitis perturbator Peringuey 1901 (BMNH drawer 184)
  • Onitis podicinus Boucomont 1932 (BMNH drawer 183)
  • Onitis pyramus Gillet 1933 (BMNH drawer 184)
  • Onitis reichei Lansberge 1875 (BMNH drawer 181)
  • Onitis retrodentatus Gillet 1932 (BMNH drawer 183)
  • Onitis robustus Boheman 1857 (BMNH drawer 180)
  • Onitis sphinx Fabricius 1775 (BMNH drawer 181)
  • Onitis spinicrus Fairmaire 1893
  • Onitis subcrenatus Kolbe 1897 (BMNH drawer 183)
  • Onitis sulcipennis Felsche 1907 (BMNH drawer 184)
  • Onitis uncinatoides Janssens 1951 (BMNH drawer 184)
  • Onitis uncinatus Klug 1855 (BMNH drawer 179)
  • Onitis upembanus Janssens 1951
  • Onitis vanderkelleni Lansberge 1886 (BMNH drawer 183)
  • Onitis viridulus Boheman 1857 (BMNH drawer 183)
  • Onitis vrydaghi Janssens 1937 (BMNH drawer 183)
  • Onitis wittei Janssens 1938 (BMNH drawer 184)
• ONTHOPHAGUS
  
  • Onthophagus abruptus d'Orbigny 1913 (BMNH drawer 265)
  • Onthophagus abyssinicus Peringuey 1901
  • Onthophagus acutus d'Orbigny 1908 (BMNH drawer 273)
  • Onthophagus aeneoniger d'Orbigny 1913
  • Onthophagus aequatus Peringuey 1901 (BMNH drawer 269)
  • Onthophagus aeruginosus Roth 1851 (BMNH drawer 256)
  • Onthophagus aethiopicus d'Orbigny 1902 (BMNH drawer 272)
  • Onthophagus africanus Lansberge 1886
  • Onthophagus ahenomicans d'Orbigny 1902
  • Onthophagus alienus Frey 1958
  • Onthophagus alluaudi d'Orbigny 1902 (BMNH drawer 261)
  • Onthophagus altidorsis d'Orbigny 1905 (BMNH drawer 268)
  • Onthophagus altilamina d'Orbigny 1905
  • Onthophagus angularis d'Orbigny 1908 (BMNH drawer 256)
  • Onthophagus antennalis Frey 1961
  • Onthophagus apiciosus d'Orbigny 1902 (BMNH drawer 266)
  • Onthophagus asperipennis d'Orbigny 1902
  • Onthophagus atroaereus d'Orbigny 1908
  • Onthophagus atrostriatus d'Orbigny 1913 (BMNH drawer 259)
  • Onthophagus basilewskyi Frey 1961
  • Onthophagus bellus d'Orbigny 1905
  • Onthophagus bequaerti d'Orbigny 1915
  • Onthophagus biconifer d'Orbigny 1905 (BMNH drawer 256)
  • Onthophagus bidens Olivier 1789 (BMNH drawer 269)
  • Onthophagus bidentifrons d'Orbigny 1902
  • Onthophagus bifidus Reiche 1849 (BMNH drawer 273)
  • Onthophagus bimarginatus d'Orbigny 1902 (BMNH drawer 266)
  • Onthophagus bisbicornis d'Orbigny 1909
  • Onthophagus bituber d'Orbigny 1904
  • Onthophagus bituberculatus Olivier 1789 (BMNH drawer 273)
  • Onthophagus brevipennis d'Orbigny 1902
  • Onthophagus callosipennis Boucomont 1930
  • Onthophagus cancer Lansberge 1886 (BMNH drawer 268)
  • Onthophagus carcharias Harold 1875 (BMNH drawer 256)
  • Onthophagus chlorophanus d'Orbigny 1902 (BMNH drawer 256)
  • Onthophagus cinctipennis Quedenfeldt 1884 (BMNH drawer 256)
  • Onthophagus clavisetis d'Orbigny 1905 (BMNH drawer 257)
  • Onthophagus cornifrons Thomson 1858 (BMNH drawer 258)
  • Onthophagus costatus d'Orbigny 1913
  • Onthophagus cupreovirens d'Orbigny 1904
  • Onthophagus cupreus Harold 1880 (BMNH drawer 264)
  • Onthophagus curtulus Frey 1961
  • Onthophagus cyaneoniger d'Orbigny 1902
  • Onthophagus cyanochlorus d'Orbigny 1902 (BMNH drawer 269)
  • Onthophagus deflexus d'Orbigny 1908 (BMNH drawer 266)
  • Onthophagus delicatus d'Orbigny 1915
  • Onthophagus densipilis d'Orbigny 1902 (BMNH drawer 270)
  • Onthophagus denticulatus d'Orbigny 1902 (BMNH drawer 268)
  • Onthophagus depilis d'Orbigny 1902
  • Onthophagus depressifrons d'Orbigny 1915
  • Onthophagus desaegeri Frey 1961
  • Onthophagus duvivieri d'Orbigny 1904 (BMNH drawer 263)
  • Onthophagus epilamprus Bates 1888 (BMNH drawer 266)
  • Onthophagus fasciolatus Boucomont 1924 (BMNH drawer 274)
  • Onthophagus filicornis Harold 1873 (BMNH drawer 275)
  • Onthophagus flaviclava d'Orbigny 1902 (BMNH drawer 273)
  • Onthophagus foveatus Frey 1957
  • Onthophagus foveicollis Boucomont 1930
  • Onthophagus fungicola d'Orbigny 1908 (BMNH drawer 262)
  • Onthophagus fuscatus d'Orbigny 1908 (BMNH drawer 261)
  • Onthophagus gaillardi d'Orbigny 1911
  • Onthophagus garambae Frey 1961
  • Onthophagus gibbifrons d'Orbigny 1902 (BMNH drawer 266)
  • Onthophagus gonopygus d'Orbigny 1902 (BMNH drawer 270)
  • Onthophagus graniceps d'Orbigny 1908
  • Onthophagus granulifer Harold 1886 (BMNH drawer 270)
  • Onthophagus helciatus Harold 1871 (BMNH drawer 266)
  • Onthophagus hemichalceus d'Orbigny 1915
  • Onthophagus herus Peringuey 1901 (BMNH drawer 264)
  • Onthophagus hirtellus Frey 1957
  • Onthophagus hulstaerti Boucomont 1932 (BMNH drawer 274)
  • Onthophagus illotus Peringuey 1900 (BMNH drawer 257)
  • Onthophagus imberbis d'Orbigny 1902 (BMNH drawer 258)
  • Onthophagus impressicollis Boheman 1860 (BMNH drawer 268)
  • Onthophagus indigus Peringuey 1901 (BMNH drawer 263)
  • Onthophagus ineptus Harold 1871 (BMNH drawer 262)
  • Onthophagus inquisitus Frey 1961
  • Onthophagus insignis Peringuey 1896 (BMNH drawer 268)
  • Onthophagus interstitialis Fahraeus 1857 (BMNH drawer 258)
  • Onthophagus janssensi Frey 1958
  • Onthophagus jugicola d'Orbigny 1902 (BMNH drawer 262)
  • Onthophagus kassaicus d'Orbigny 1908
  • Onthophagus katangensis Frey 1961
  • Onthophagus kulzeri Frey 1958
  • Onthophagus lacustris Harold 1877 (BMNH drawer 261)
  • Onthophagus laeviceps d'Orbigny 1902 (BMNH drawer 267)
  • Onthophagus lamellicornis Frey 1953 (BMNH drawer 274)
  • Onthophagus laminicornis Lansberge 1886
  • Onthophagus latestriatus d'Orbigny 1908 (BMNH drawer 258)
  • Onthophagus lemekensis d'Orbigny 1915 (BMNH drawer 274)
  • Onthophagus lognegranus Frey 1957
  • Onthophagus lutaticollis d'Orbigny 1907 (BMNH drawer 268)
  • Onthophagus macrothorax d'Orbigny 1902
  • Onthophagus marginifer Frey 1953
  • Onthophagus masaicus d'Orbigny 1905 (BMNH drawer 256)
  • Onthophagus minax d'Orbigny 1904 (BMNH drawer 263)
  • Onthophagus misellus d'Orbigny 1907 (BMNH drawer 256)
  • Onthophagus mixticeps d'Orbigny 1905 (BMNH drawer 268)
  • Onthophagus mocquerysi d'Orbigny 1902 (BMNH drawer 267)
  • Onthophagus mucronatus Thomson 1858 (BMNH drawer 269)
  • Onthophagus musculus Frey 1963
  • Onthophagus naevius d'Orbigny 1913 (BMNH drawer 269)
  • Onthophagus nitidifrons d'Orbigny 1905
  • Onthophagus ocellidorsis d'Orbigny 1915 (BMNH drawer 275)
  • Onthophagus ochropygus d'Orbigny 1902 (BMNH drawer 272)
  • Onthophagus ornatulus d'Orbigny 1908
  • Onthophagus orthocerus Thomson 1958 (BMNH drawer 264)
  • Onthophagus parcepictus d'Orbigny 1908 (BMNH drawer 263)
  • Onthophagus parumnotatus Fahraeus 1857 (BMNH drawer 263)
  • Onthophagus paulianellus Frey 1959
  • Onthophagus phanaeomorphus Janssens 1954 (BMNH drawer 274)
  • Onthophagus picatus d'Orbigny 1902 (BMNH drawer 261)
  • Onthophagus pictisternum d'Orbigny 1902
  • Onthophagus picturatus d'Orbigny 1908
  • Onthophagus pinguis Gerstaecker 1871 (BMNH drawer 266)
  • Onthophagus planaticeps d'Orbigny 1913 (BMNH drawer 264)
  • Onthophagus portentosus Felsche 1907 (BMNH drawer 256)
  • Onthophagus praestans Peringuey 1900 (BMNH drawer 269)
  • Onthophagus profanus Frey 1958
  • Onthophagus proximus d'Orbigny 1908 (BMNH drawer 268)
  • Onthophagus pseudoaeneus d'Orbigny 1908 (BMNH drawer 263)
  • Onthophagus pseudobidens d'Orbigny 1915
  • Onthophagus pseudopilosus Frey 1958
  • Onthophagus pugionatus Fahraeus 1857
  • Onthophagus pulchellus d'Orbigny 1905
  • Onthophagus pullatus d'Orbigny 1905
  • Onthophagus pullus Roth (BMNH drawer 262)
  • Onthophagus quadrinodosus Fahraeus 1857 (BMNH drawer 264)
  • Onthophagus rectestriatus d'Orbigny 1915
  • Onthophagus reticulatus d'Orbigny 1802 (BMNH drawer 258)
  • Onthophagus reticuliger Frey 1960
  • Onthophagus rhynchophorus Peringuey 1904 (BMNH drawer 266)
  • Onthophagus rosettae Frey 1958
  • Onthophagus rubricatus d'Orbigny 1902 (BMNH drawer 266)
  • Onthophagus rufaticollis d'Orbigny 1904 (BMNH drawer 269)
  • Onthophagus rufobasalis Fairmaire 1887 (BMNH drawer 269)
  • Onthophagus rufonotatus d'Orbigny 1902
  • Onthophagus rufostillans d'Orbigny 1907 (BMNH drawer 269)
  • Onthophagus sansibaricus Harold 1878 (BMNH drawer 263)
  • Onthophagus schoutedeni Bocuomont 1920 (BMNH drawer 274)
  • Onthophagus sellatus Klug 1845 (BMNH drawer 231)
  • Onthophagus semicroceus d'Orbigney
  • Onthophagus semivirescens d'Orbigny 1902 (BMNH drawer 267)
  • Onthophagus sericeicollis Boucomont 1928 (BMNH drawer 275)
  • Onthophagus simoni d'Orbigny 1902 (BMNH drawer 256)
  • Onthophagus simplex Raffray 1877 (BMNH drawer 262)
  • Onthophagus simpliciceps d'Orbigny 1905 (BMNH drawer 264)
  • Onthophagus splendidus Boucomont 1932
  • Onthophagus subdivisus d'Orbigny 1908
  • Onthophagus subulifer d'Orbigny 1908 (BMNH drawer 270)
  • Onthophagus sulcipennis d'Orbigny 1902 (BMNH drawer 261)
  • Onthophagus tenuigraniger d'Orbigny 1913 (BMNH drawer 259)
  • Onthophagus tibialis Frey 1963 (BMNH drawer 274)
  • Onthophagus tigrinus d'Orbigny 1908 (BMNH drawer 257)
  • Onthophagus tonsus d'Orbigny 1902 (BMNH drawer 262)
  • Onthophagus trapezicornis d'Orbigny 1902 (BMNH drawer 268)
  • Onthophagus trichopygus d'Orbigny 1905 (BMNH drawer 264)
  • Onthophagus tridentipes d'Orbigny 1902
  • Onthophagus triimpressus d'Orbigny 1905 (BMNH drawer 261)
  • Onthophagus triundulatus Balthasar 1964
  • Onthophagus tschoffeni d'Orbigny 1904
  • Onthophagus tschuapae Balthasar 1964
  • Onthophagus uelensis Frey 1960
  • Onthophagus umbratus d'Orbigny 1902 (BMNH drawer 261)
  • Onthophagus vaneyeni Frey 1960
  • Onthophagus variatus d'Orbigny 1902
  • Onthophagus variegranosus d'Orbigny 1905 (BMNH drawer 263)
  • Onthophagus velliger d'Orbigny 1907
  • Onthophagus vinctoides Frey 1957 (BMNH drawer 258)
  • Onthophagus vinctus Erichson 1843 (BMNH drawer 258)
  • Onthophagus viviensis d'Orbigny 1905
  • Onthophagus vultuosus d'Orbigny 1902 (BMNH drawer 267)
  • Onthophagus wittei Frey 1958
  • Onthophaus pallidus d'Orbigny
• PACHYLOMERUS
  
  • Pachylomerus femoralis Kirby 1828 (BMNH drawer 24)
• PARAPHYTUS
  
  • Paraphytus ritsemae Harold 1880
• PEDARIA
  
  • Pedaria criberrima Waterhouse 1890 (BMNH drawer 72)
  • Pedaria hanae Balthasar 1963
  • Pedaria intermedia Boucomont 1922 (BMNH drawer 72)
  • Pedaria ovata Boucomont 1922
  • Pedaria spinosa Boucomont 1922 (BMNH drawer 72)
  • Pedaria tuberculigera Waterhouse 1890 (BMNH drawer 72)
  • Pedaria wittei Janssens 1939
• PHALOPS
  
  • Phalops batesi Harold 1867 (BMNH drawer 206)
  • Phalops boschas Klug 1855 (BMNH drawer 208)
  • Phalops coeruleatus d'Orbigny 1902
  • Phalops prasinus Erichson 1843 (BMNH drawer 208)
  • Phalops vanellus Lansberge 1883 (BMNH drawer 207)
• PROAGODERUS
  
  • Proagoderus armicollis d'Orbigny 1904 (BMNH drawer 216)
  • Proagoderus auratus Fabricius 1801 (BMNH drawer 217)
  • Proagoderus brucei Reiche 1849 (BMNH drawer 218)
  • Proagoderus ceratotherium Janssens 1953
  • Proagoderus colmanti d'Orbigny 1908
  • Proagoderus confossus d'Orbigny 1915 (BMNH drawer 216)
  • Proagoderus cuspidatus d'Orbigny 1913
  • Proagoderus elgoni d'Orbigny 1915 (BMNH drawer 215)
  • Proagoderus extensus Harold 1878 (BMNH drawer 216)
  • Proagoderus fastosus d'Orbigny 1913
  • Proagoderus fossidorsis d'Orbigny 1905
  • Proagoderus gemmatus Peringuey 1901 (BMNH drawer 218)
  • Proagoderus gibbiramus d'Orbigny 1902 (BMNH drawer 212)
  • Proagoderus harpax Fabricius 1801 (BMNH drawer 217)
  • Proagoderus katualensis Quedenfeldt 1888
  • Proagoderus lallieri d'Orbigny 1902 (BMNH drawer 216)
  • Proagoderus lallierianus Janssens 1953
  • Proagoderus lujendae Bates 1888 (BMNH drawer 216)
  • Proagoderus monochromus d'Orbigny 1913
  • Proagoderus multicornis d'Orbigny 1908 (BMNH drawer 215)
  • Proagoderus overlaeti Janssens 1953
  • Proagoderus panoplus Bates 1888 (BMNH drawer 215)
  • Proagoderus pegesimallus Marcus 1917
  • Proagoderus praedentatus d'Orbigny 1904
  • Proagoderus quadrispinosus d'Orbigny 1908
  • Proagoderus quadrituber d'Orbigny 1908 (BMNH drawer 217)
  • Proagoderus rangifer Klug 1855 (BMNH drawer 212)
  • Proagoderus rarus Guerin 1829 (BMNH drawer 212)
  • Proagoderus semiiris Thomson 1858 (BMNH drawer 218)
  • Proagoderus speculicollis Quedenfeldt 1884 (BMNH drawer 217)
  • Proagoderus subextensus Kolbe 1895 (BMNH drawer 216)
  • Proagoderus superbus d'Orbigny 1904
  • Proagoderus taymansi d'Orbigny 1908
  • Proagoderus trianguliceps d'Orbigny 1905 (BMNH drawer 218)
  • Proagoderus unidens Quedenfeldt 1884 (BMNH drawer 249)
• PSEUDOCOPRIS
  
  • Pseudocopris tubericollis Felsche 1910 (BMNH drawer 142)
• PSEUDOPEDARIA
  
  • Pseudopedaria grossa Thomson 1858 (BMNH drawer 142)
• PSEUDOSAPROECIUS
  
  • Pseudosaproecius cylindroides d'Orbigny 1908 (BMNH drawer 70)
  • Pseudosaproecius dicerus d'Orbigny 1915
  • Pseudosaproecius mirepunctatus d'Orbigny 1905
  • Pseudosaproecius ocellatus Frey 1958
  • Pseudosaproecius splendens Frey 1961
  • Pseudosaproecius validicornis Quedenfeldt 1884 (BMNH drawer 70)
• SAPROECIUS
  
  • Saproecius hirsutus Boucomont 1926
• SCAPTOCNEMIS
  
  • Scaptocnemis segregis Peringuey 1901 (BMNH drawer 187)
• SCARABAEOLUS
  
  • Scarabaeolus ebenus Klug 1855 (BMNH drawer 3)
• SCARABAEUS
  
  • Scarabaeus goryi Castelnau 1840 (BMNH drawer 11)
  • Scarabaeus kwiluensis Janssens 1940 (BMNH drawer 2)
  • Scarabaeus palemo Olivier 1789 (BMNH drawer 1)
  • Scarabaeus poggei Waterhouse 1890 (BMNH drawer 8)
  • Scarabaeus sulcipennis Quedenfeldt 1888 (BMNH drawer 8)
• SCELIAGES
  
  • Sceliages augias Gillet 1908 (BMNH drawer 27.1)
• SISYPHUS
  
  • Sisyphus costatus Thunberg 1818 (BMNH drawer 40)
  • Sisyphus crispatus Gory 1833 (BMNH drawer 40)
  • Sisyphus transvaalensis Peringuey 1908 (BMNH drawer 40 & 41)
• STIPTOTARSUS
  
  • Stiptotarsus tersus Balthasar 1963
• STRANDIUS
  
  • Strandius obliquus Olivier 1789 (BMNH drawer 257)
• TINIOCELLUS
  
  • Tiniocellus collarti Janssens 1939
  • Tiniocellus panthera Boucomont 1921
  • Tiniocellus spinipes Roth 1851 (BMNH drawer 190)
• TOMOGONUS
  
  • Tomogonus crassus d'Orbigny 1904
• TRAGISCUS
  
  • Tragiscus dimidiatus Klug 1855 (BMNH drawer 194)
• TRICHONTHOPHAGUS
  
  • Trichonthophagus corniculatus Reiche 1849 (BMNH drawer 259)
  • Trichonthophagus juvencus Klug 1835 (BMNH drawer 259)
  • Trichonthophagus quadricuspis d'Orbigny 1908 (BMNH drawer 259)
• WALTERANTUS
  
  • Walterantus rufoclypeatus Frey 1960