adapted from Monaghan et al., (2007)
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Scarab Classification: Amphistomus; Anachalcos; Anonthobium; Apotolamprus; Aptenocanthon; Arachnodes; Arachnodes splendidus; Ateuchus; Bdelyropsis; Bubas; Caccobius; Canthidium; Canthon; Canthonosoma; Catharsius; Cephalodesmius; Circellium; Cleptocaccobius; Copris; Coproecus; Coprophanaeus; Coptodactyla; Coptorhina; Deltochilum; Demarziella; Diabroctis; Dichotomius; Dicranocara; Digitonthophagus; Diorygopyx; Drepanocerus; Ennearabdus; Epirinus; Eucranium; Eudinopus; Euoniticellus; Eurysternus; Glyphoderus; Gromphas; Gymnopleurus; Hansreia; Helictopleurus; Heliocopris; Heteronitis; Ignambia; Lepanus; Liatongus; Megathoposoma; Metacatharsius; Milichus; Monoplistes; Neosisyphus; Odontoloma; Odontoloma pusillum; Oniticellus; Onitis; Ontherus; Onthobium; Onthophagus; Pachylomerus; Pachysoma; Panelus; Paragymnopleurus; Paronthobium; Pedaria; Phalops; Phanaeus; Proagoderus; Pseudignambia; Pseudonthobium; Saphobius; Sarophorus; Scarabaeus; Sceliages; Scybalocanthon; Scybalophagus; Sisyphus; Sphaerocanthon; Temnoplectron; Tiniocellus; Tragiscus; Trichillum; Uroxys
