Dung beetles found in Tanzania

12 species of dung beetles are known to occur in Tanzania none of which are endemic. If you are aware of other species not included here please e-mail christopherborrow@gmail.com.

ALLOGYMNOPLEURUS
- Allogymnopleurus aeneus Harold, 1875 (BMNH drawer 34)
- Allogymnopleurus chloris Klug, 1855 (BMNH drawer 34)
- Allogymnopleurus histrio Balthasar, 1963 (BMNH drawer 34.1)
- Allogymnopleurus jeanneli Garreta, 1914 (BMNH drawer 34)
- Allogymnopleurus thalassinus Klug, 1855 (BMNH drawer 34)
- Allogymnopleurus umbrinus Gerstaecker, 1871 (BMNH drawer 34.1)
ALLOSCELUS
- Alloscelus paradoxus Boucomont, 1923 (BMNH drawer 43)
ANACHALCOS
- Anachalcos aurescens Bates, 1888 (BMNH drawer 64)
- Anachalcos convexus Boheman, 1857 (BMNH drawer 63)
- Anachalcos cupreus Fabricius, 1775 (BMNH drawer 63)
- Anachalcos procerus Gerstaecker, 1874 (BMNH drawer 64)
- Anachalcos schultzi Kolbe, 1904 (BMNH drawer 63)
APTYCHONITIS
- Aptychonitis anomalus Gestro, 1895 (BMNH drawer 172)
CACCOPHILUS
- Caccophilus bicornutus Frey, 1955
- Caccophilus convexifrons Raffray, 1877 (BMNH drawer 203)
- Caccophilus ferrugineus Fahraeus, 1857 (BMNH drawer 201)
- Caccophilus obtusus Fahraeus, 1857 (BMNH drawer 201)
- Caccophilus schaedlei d'Orbigny, 1902 (BMNH drawer 203)
- Caccophilus semiluteus d'Orbigny, 1905
- Caccophilus viridicollis Fahraeus, 1857 (BMNH drawer 203)
CATHARSIUS
- Catharsius achates Olivier, 1789 (BMNH drawer 115)
- Catharsius alpheus Gillet, 1933 (BMNH drawer 116)
- Catharsius dux Harold, 1878 (BMNH drawer 113)
- Catharsius erechtheus Felsche, 1907
- Catharsius gorilla Thomson, 1858 (BMNH drawer 113)
- Catharsius hertli Ferreira, 1964
- Catharsius kolbei Felsche, 1907
- Catharsius neptunus Kolbe, 1893 (BMNH drawer 117)
- Catharsius oedipus Felsche, 1907 (BMNH drawer 116)
- Catharsius pandion Harold, 1877
- Catharsius philus Kolbe, 1893 (BMNH drawer 118)
- Catharsius platicornis Ferreira, 1960
- Catharsius satyrus Kolbe, 1893 (BMNH drawer 114)
- Catharsius scopas Felsche, 1907
- Catharsius sesostris Waterhouse, 1888 (BMNH drawer 118)
- Catharsius spectabilis Gillet, 1918 (BMNH drawer 113)
- Catharsius stuhlmanni Kolbe, 1893 (BMNH drawer 116)
- Catharsius tricornutus Degeer, 1778 (BMNH drawer 117)
CHIRONITIS
- Chironitis benningseni Felsche, 1907 (BMNH drawer 172)
- Chironitis lohsei Balthasar, 1968
COPRIS
- Copris armiger Gillet, 1910 (BMNH drawer 137)
- Copris atropolitus Gillet, 1910 (BMNH drawer 133)
- Copris bootes Klug, 1855 (BMNH drawer 132)
- Copris brittoni Ferreira, 1962 (BMNH drawer 136)
- Copris cribricollis Gillet, 1910 (BMNH drawer 133)
- Copris diversus Waterhouse, 1891 (BMNH drawer 137)
- Copris elpheneroides Felsche, 1910 (BMNH drawer 131)
- Copris elphenor Klug, 1855 (BMNH drawer 131)
- Copris evanidus Klug, 1855 (BMNH drawer 136)
- Copris gilleti Kolbe, 1907
- Copris graueri Gillet, 1910
- Copris harrisi Waterhouse, 1891 (BMNH drawer 134)
- Copris humilis Gillet, 1908 (BMNH drawer 136)
- Copris hybridus Gillet, 1910
- Copris insidiosus Peringuey, 1901 (BMNH drawer 137)
- Copris integer Reische, 1847 (BMNH drawer 132)
- Copris interioris Kolbe, 1897 (BMNH drawer 134)
- Copris macer Peringuey, 1901 (BMNH drawer 137)
- Copris nepos Gillet, 1908 (BMNH drawer 136)
- Copris obesus Boheman, 1857 (BMNH drawer 137)
- Copris orion Klug, 1835 (BMNH drawer 136)
- Copris orphanus Guerin, 1847 (BMNH drawer 137)
- Copris pesudolunaris Ferreira, 1969
- Copris pseudobootes Ferreira, 1962
- Copris usambaricus Gillet, 1908 (BMNH drawer 137)
COPTORHINA
- Coptorhina bicolor Ancey, 1882 (BMNH drawer 71)
- Coptorhina davidi Janssens, 1939
- Coptorhina nitidipennis Boheman, 1857 (BMNH drawer 71)
DIASTELLOPALPUS
- Diastellopalpus fulleborni Kolbe, (BMNH drawer 210)
- Diastellopalpus infernalis Lansberge, 1887 (BMNH drawer 209)
- Diastellopalpus johnstoni Waterhouse, 1885 (BMNH drawer 210)
- Diastellopalpus laevibasis d'Orbigny, 1902 (BMNH drawer 209)
- Diastellopalpus lamellicollis Quedenfeldt, 1884 (BMNH drawer 209)
- Diastellopalpus metriogonus Marcus, 1914 (BMNH drawer 210)
- Diastellopalpus monapoides Bates, 1888
- Diastellopalpus quinquedens Bates, 1888 (BMNH drawer 209)
- Diastellopalpus thomsoni Bates, 1888 (BMNH drawer 209)
DIGITONTHOPHAGUS
- Digitonthophagus gazella Fabricius, 1787 (BMNH drawer 221)
DREPANOCERUS
- Drepanocerus abyssinicus Roth, 1851 (BMNH drawer 186)
- Drepanocerus bechynei Janssens, 1953
- Drepanocerus caelatus Gerstaecker, 1871 (BMNH drawer 186)
- Drepanocerus laticollis Fahraeus, 1857 (BMNH drawer 186)
- Drepanocerus striatus Boucomont, 1921 (BMNH drawer 186)
- Drepanocerus sulcicollis Castelnau, 1840 (BMNH drawer 186)
EPIONITIS
- Epionitis tarsatus Balthasar, 1942
EUONITICELLUS
- Euoniticellus inaequalis Reiche, 1850 (BMNH drawer 190)
- Euoniticellus intermedius Reiche, 1849 (BMNH drawer 188)
- Euoniticellus nasicornis Reiche, 1849 (BMNH drawer 190)
- Euoniticellus parvus Kraatz, 1895 (BMNH drawer 188)
- Euoniticellus triangulatus Harold, 1873 (BMNH drawer 190)
EUONTHOPHAGUS
- Euonthophagus carbonarius Klug, 1855 (BMNH drawer 220)
EUSAPROECIUS
- Eusaproecius magnigena d'Orbigny, 1913 (BMNH drawer 70)
FURCONTHOPHAGUS
- Furconthophagus variegatus Fabricius, 1798 (BMNH drawer 231)
GARRETA
- Garreta lugens Fairmaire, 1891 (BMNH drawer 33)
- Garreta malleolus Kolbe, 1895 (BMNH drawer 33)
GYMNOPLEURUS
- Gymnopleurus fulgidus Olivier, 1789 (BMNH drawer 36)
- Gymnopleurus jacksoni Waterhouse, 1890 (BMNH drawer 38.1)
- Gymnopleurus laevicollis Castelnau, 1840 (BMNH drawer 36)
- Gymnopleurus sericeifrons Fairmaire, 1887 (BMNH drawer 37)
GYRONOTUS
- Gyronotus dispar Felsche, 1911 (BMNH drawer 65)
- Gyronotus fimetarius Kolbe, 1894 (BMNH drawer 65)
HELIOCOPRIS
- Heliocopris alatus Felsche, 1910 (BMNH drawer 86)
- Heliocopris anadematus Gillet, 1908
- Heliocopris biimpressus Kolbe, 1893
- Heliocopris colossus Bates, 1868 (BMNH drawer 79)
- Heliocopris densissa Roth, 1851 (BMNH drawer 86)
- Heliocopris dolosus Janssens, 1940
- Heliocopris erycoides Felsche, 1907 (BMNH drawer 85)
- Heliocopris felschei Kolbe, 1904
- Heliocopris hunteri Waterhouse, 1891 (BMNH drawer 87)
- Heliocopris japetus Klug, 1855 (BMNH drawer 80)
- Heliocopris myrmidon Kolbe, 1893 (BMNH drawer 84)
- Heliocopris neptuniformis Felsche, 1907 (BMNH drawer 84)
HETEROCLITOPUS
- Heteroclitopus foveatus Boucomont, 1923 (BMNH drawer 70)
- Heteroclitopus herteli Ferreira, 1969
HETERONITIS
- Heteronitis castelnaui Harold, 1862 (BMNH drawer 174)
HYALONTHOPHAGUS
- Hyalonthophagus alcyon Klug, 1855 (BMNH drawer 217)
- Hyalonthophagus virens d'Orbigny, 1913 (BMNH drawer 217)
- Hyalonthophagus viridiceps d'Orbigny, 1910
KHEPER
- Kheper aegyptiorum Latreille, 1827 (BMNH drawer 16)
- Kheper aeratus Gerstaecker, 1871 (BMNH drawer 19)
- Kheper cupreus Castelnau, 1840 (BMNH drawer 17)
- Kheper laevistriatus Fairmaire, 1893 (BMNH drawer 19)
- Kheper platynotus Bates, 1888 (BMNH drawer 22)
- Kheper porosus Bates, 1888 (BMNH drawer 19)
- Kheper prodigiosus Erichson, 1843 (BMNH drawer 21)
- Kheper pustulosus Gerstaecker, 1871 (BMNH drawer 18)
- Kheper venerabilis Harold, 1871 (BMNH drawer 17)
LIATONGUS
- Liatongus amitinus Boucomont, 1921 (BMNH drawer 193)
- Liatongus aterrimus Boucomont, 1921
- Liatongus militaris Castelnau, 1840 (BMNH drawer 194)
- Liatongus schoutedeni Boucomont, 1920
LITOCOPRIS
- Litocopris punctiventris Waterhouse, 1891 (BMNH drawer 137)
MADAPHACOSOMA
- Madaphacosoma basilewskyi Balthasar, 1960
METACATHARSIUS
- Metacatharsius abortivus Fairmaire, 1891 (BMNH drawer 121)
- Metacatharsius nubiensis Harold, 1878 (BMNH drawer 121)
- Metacatharsius opacus Waterhouse, 1891 (BMNH drawer 120)
- Metacatharsius seminulum Balthasar, 1940
- Metacatharsius simulator Balthasar, 1939
- Metacatharsius useramus Kolbe, 1893
MILICHUS
- Milichus jugatus d'Orbigny, 1907
- Milichus picticollis Gerstaecker, 1871 (BMNH drawer 204)
- Milichus rhodesianus Peringuey, 1904 (BMNH drawer 204)
NEOSISYPHUS
- Neosisyphus armatus Gory, 1833 (BMNH drawer 42.3 & 42.4)
- Neosisyphus spinipes Thunberg, 1818 (BMNH drawer 42.3)
- Neosisyphus tibialis Raffray, 1877 (BMNH drawer 42.3)
ONITICELLUS
- Oniticellus egregius Klug, 1855 (BMNH drawer 191)
- Oniticellus pictus Hausmann, 1807 (BMNH drawer 191)
- Oniticellus planatus Castelnau, 1840 (BMNH drawer 191)
- Oniticellus pseudoplanatus Balthasar, 1964
ONITIS
- Onitis adelphus Kolbe, 1907 (BMNH drawer 180)
- Onitis alexis Klug, 1835 (BMNH drawer 181)
- Onitis anthracinus Felsche, 1907 (BMNH drawer 185)
- Onitis consaguineus Felsche, 1911
- Onitis coxalis Gillet, 1918 (BMNH drawer 185)
- Onitis deceptor Peringuey, 1901 (BMNH drawer 179)
- Onitis dimidiatus Felsche, 1907
- Onitis fulmineus Janssens, 1943
- Onitis inflaticollis Felsche, 1907 (BMNH drawer 104)
- Onitis laminifer Gillet, 1909
- Onitis lobipes Felsche, 1907 (BMNH drawer 185)
- Onitis lycophron Klug, 1855 (BMNH drawer 180)
- Onitis malleatus Janssens, 1937
- Onitis mendax Gillet, 1918 (BMNH drawer 185)
- Onitis meyeri Kolbe, 1891
- Onitis parvulus Balthasar, 1963 (BMNH drawer 185)
- Onitis robustus Boheman, 1857 (BMNH drawer 180)
- Onitis setosus Lansberge, 1875 (BMNH drawer 184)
- Onitis similis Felsche, 1911
- Onitis sphinx Fabricius, 1775 (BMNH drawer 181)
- Onitis spinicrus Fairmaire, 1893
- Onitis subcrenatus Kolbe, 1897 (BMNH drawer 183)
- Onitis sulcipennis Felsche, 1907 (BMNH drawer 184)
- Onitis uncinatus Klug, 1855 (BMNH drawer 179)
- Onitis viridulus Boheman, 1857 (BMNH drawer 183)
- Onitis westermanni Lansberge, 1886 (BMNH drawer 184)
ONTHOPHAGUS
- Onthophagus abyssinicus Peringuey, 1901
- Onthophagus aeruginosus Roth, 1851 (BMNH drawer 256)
- Onthophagus albicomus d'Orbigny, 1908 (BMNH drawer 263)
- Onthophagus amplipennis d'Orbigny, 1905
- Onthophagus apiciosus d'Orbigny, 1902 (BMNH drawer 266)
- Onthophagus areolatus d'Orbigny, 1905
- Onthophagus asperatus d'Orbigny, 1905 (BMNH drawer 264)
- Onthophagus aspernatus d'Orbigny, 1907
- Onthophagus asperrimus d'Orbigny, 1902 (BMNH drawer 264)
- Onthophagus astigma d'Orbigny, 1904
- Onthophagus atrofasciatus d'Orbigny, 1905 (BMNH drawer 270)
- Onthophagus bellus d'Orbigny, 1905
- Onthophagus bicristiger d'Orbigny, 1913
- Onthophagus bidens Olivier, 1789 (BMNH drawer 269)
- Onthophagus bifidicornis d'Orbigny, 1902
- Onthophagus bifrons d'Orbigny, 1905
- Onthophagus bituberculatus Olivier, 1789 (BMNH drawer 273)
- Onthophagus carinicollis Raffray, 1877 (BMNH drawer 256)
- Onthophagus carinidorsis d'Orbigny, 1908
- Onthophagus chlorophanus d'Orbigny, 1902 (BMNH drawer 256)
- Onthophagus clitellarius d'Orbigny, 1908
- Onthophagus clusifrons d'Orbigny, 1905
- Onthophagus comatulus d'Orbigny, 1905 (BMNH drawer 263)
- Onthophagus conradsi Balthasar, 1937
- Onthophagus contiguicornis d'Orbigny, 1913 (BMNH drawer 270)
- Onthophagus cornutus Ferreira, 1971
- Onthophagus costilatus d'Orbigny, 1908
- Onthophagus creber d'Orbigny, 1902
- Onthophagus cristatus d'Orbigny, 1905
- Onthophagus debilis d'Orbigny, 1905
- Onthophagus densatus Frey, 1963
- Onthophagus dicax Balthasar, 1966
- Onthophagus dicella Bates, 1888 (BMNH drawer 264)
- Onthophagus dispar Peringuey, 1901 (BMNH drawer 264)
- Onthophagus diversus Reiche, 1847 (BMNH drawer 269)
- Onthophagus dlabolai Balthasar, 1964
- Onthophagus ebenicolor d'Orbigny, 1902 (BMNH drawer 266)
- Onthophagus ebenus Peringuey, 1888 (BMNH drawer 270)
- Onthophagus excisiceps d'Orbigny, 1902 (BMNH drawer 270)
- Onthophagus falcarius Balthasar, 1966
- Onthophagus fallax d'Orbigny, 1913 (BMNH drawer 273)
- Onthophagus filicornis Harold, 1873 (BMNH drawer 275)
- Onthophagus fimetarius Roth, 1851 (BMNH drawer 268)
- Onthophagus fossibasis d'Orbigny, 1908
- Onthophagus fossulatus d'Orbigny, 1908
- Onthophagus fulvocinctus d'Orbigny, 1902
- Onthophagus fuscivestis d'Orbigny, 1902
- Onthophagus haafi Frey, 1963
- Onthophagus herus Peringuey, 1901 (BMNH drawer 264)
- Onthophagus impressicollis Boheman, 1860 (BMNH drawer 268)
- Onthophagus impurus Harold, 1868 (BMNH drawer 266)
- Onthophagus insignis Peringuey, 1896 (BMNH drawer 268)
- Onthophagus jugicola d'Orbigny, 1902 (BMNH drawer 262)
- Onthophagus juncticornis d'Orbigny, 1908
- Onthophagus laceratus Gerstaecker, 1871 (BMNH drawer 273)
- Onthophagus lacustris Harold, 1877 (BMNH drawer 261)
- Onthophagus laevidorsis d'Orbigny, 1907
- Onthophagus lagnyi Paulian, 1937
- Onthophagus lamelliger Gerstaecker, 1871 (BMNH drawer 270)
- Onthophagus laminidorsis d'Orbigny, 1902 (BMNH drawer 266)
- Onthophagus latestriatus d'Orbigny, 1908 (BMNH drawer 258)
- Onthophagus latevittatus d'Orbigny, 1905
- Onthophagus latissimus d'Orbigny, 1902
- Onthophagus leroyi d'Orbigny, 1902 (BMNH drawer 268)
- Onthophagus liopterus Harold, 1880 (BMNH drawer 261)
- Onthophagus longiceps d'Orbigny, 1904 (BMNH drawer 264)
- Onthophagus meruanus d'Orbigny, 1908
- Onthophagus miscellus d'Orbigny, 1905 (BMNH drawer 269)
- Onthophagus morosus Gerstaecker, 1871
- Onthophagus nigrinus Paulian, 1937
- Onthophagus nigriventris d'Orbigny, 1905 (BMNH drawer 257)
- Onthophagus obtegens Harold, 1886
- Onthophagus oldius Balthasar, 1964
- Onthophagus omostigma d'Orbigny, 1902 (BMNH drawer 273)
- Onthophagus parcepictus d'Orbigny, 1908 (BMNH drawer 263)
- Onthophagus parumnotatus Fahraeus, 1857 (BMNH drawer 263)
- Onthophagus parvifrons d'Orbigny, 1907 (BMNH drawer 267)
- Onthophagus perniger Boucomont, 1930 (BMNH drawer 275)
- Onthophagus pictipodex d'Orbigny, 1902 (BMNH drawer 259)
- Onthophagus plebejus Klug, 1855 (BMNH drawer 257)
- Onthophagus plicatifrons d'Orbigny, 1908
- Onthophagus polyedrus d'Orbigny, 1905 (BMNH drawer 265)
- Onthophagus polyodon d'Orbigny, 1913 (BMNH drawer 261)
- Onthophagus polystigma d'Orbigny, 1902 (BMNH drawer 262)
- Onthophagus portentosus Felsche, 1907 (BMNH drawer 256)
- Onthophagus protuberans Frey, 1955 (BMNH drawer 274)
- Onthophagus pseudofimetarius Balthasar, 1946
- Onthophagus pugionatus Fahraeus, 1857
- Onthophagus pullus Roth, (BMNH drawer 262)
- Onthophagus quadrilunatus d'Orbigny, 1902
- Onthophagus quadrimaculatus Raffray, 1877 (BMNH drawer 256)
- Onthophagus rubefactus d'Orbigny, 1913
- Onthophagus rubenticollis d'Orbigny, 1902
- Onthophagus rufobasalis Fairmaire, 1887 (BMNH drawer 269)
- Onthophagus rufocastaneus Fairmaire, 1887
- Onthophagus rugidorsis d'Orbingy, 1913
- Onthophagus sansibaricus Harold, 1878 (BMNH drawer 263)
- Onthophagus sculptilis Gerstaecker, 1871
- Onthophagus simoni d'Orbigny, 1902 (BMNH drawer 256)
- Onthophagus simplex Raffray, 1877 (BMNH drawer 262)
- Onthophagus simpliciceps d'Orbigny, 1905 (BMNH drawer 264)
- Onthophagus snoflaki Balthasar, 1943
- Onthophagus spurcatus d'Orbigny, 1905 (BMNH drawer 262)
- Onthophagus stigmosus d'Orbigny, 1902 (BMNH drawer 262)
- Onthophagus stuhlmanni d'Orbigny, 1908 (BMNH drawer 256)
- Onthophagus subhumeralis d'Orbigny, 1902
- Onthophagus subrugosus d'Orbigny, 1905
- Onthophagus sutiliceps d'Orbigny, 1902 (BMNH drawer 267)
- Onthophagus taboranus d'Orbigny, 1902
- Onthophagus tanganus d'Orbigny, 1913
- Onthophagus teitanicus d'Orbigny, 1902 (BMNH drawer 263)
- Onthophagus tenax Balthasar, 1964
- Onthophagus tonsus d'Orbigny, 1902 (BMNH drawer 262)
- Onthophagus tricariniger d'Orbigny, 1902 (BMNH drawer 258)
- Onthophagus ukerewensis Balthasar, 1937 (BMNH drawer 275)
- Onthophagus umbilicatus d'Orbigny, 1908
- Onthophagus undaticeps d'Orbigny, 1902 (BMNH drawer 266)
- Onthophagus unicarina d'Orbigny, 1904 (BMNH drawer 264)
- Onthophagus urus Boucomont, 1925
- Onthophagus verrucosus d'Orbigny, 1902
- Onthophagus vinctus Erichson, 1843 (BMNH drawer 258)
- Onthophagus xanthopygus d'Orbigny, 1908
PACHYLOMERUS
- Pachylomerus femoralis Kirby, 1828 (BMNH drawer 24)
PEDARIA
- Pedaria armata Raffray, 1877 (BMNH drawer 72)
- Pedaria insularis Kolbe, 1914
- Pedaria intermedia Boucomont, 1922 (BMNH drawer 72)
- Pedaria segregis Peringuey, 1901 (BMNH drawer 72)
- Pedaria taylori Waterhouse, 1890 (BMNH drawer 72)
- Pedaria tenebrosa Balthasar, 1937
PHALOPS
- Phalops ardea Klug, 1855 (BMNH drawer 207)
- Phalops boschas Klug, 1855 (BMNH drawer 208)
- Phalops flavocinctus Klug, 1855 (BMNH drawer 207)
- Phalops glauningi Kuntzen, 1913
- Phalops guttulatus Fairmaire, 1887
- Phalops laminifrons Fairmaire, 1882 (BMNH drawer 206)
- Phalops lutatus d'Orbigny, 1902 (BMNH drawer 208)
- Phalops sinuaticeps d'Orbigny, 1902 (BMNH drawer 206)
- Phalops smaragdinus Harold, 1875 (BMNH drawer 206)
- Phalops trifurcus d'Orbigny, 1913 (BMNH drawer 207)
- Phalops tuberosus d'Orbigny, 1905
PINACOTARSUS
- Pinacotarsus discolor Boucomont, 1923
- Pinacotarsus proximus Ferreira, 1969
PROAGODERUS
- Proagoderus amblychromatus Marcus, 1917
- Proagoderus atriclaviger d'Orbigny, 1905 (BMNH drawer 216)
- Proagoderus biarmatus d'Orbigny, 1908
- Proagoderus bicallosus Klug, 1855 (BMNH drawer 217)
- Proagoderus chrysopes Bates, 1888 (BMNH drawer 218)
- Proagoderus dives Harold, 1877 (BMNH drawer 218)
- Proagoderus extensus Harold, 1878 (BMNH drawer 216)
- Proagoderus fossidorsis d'Orbigny, 1905
- Proagoderus furciramus d'Orbigny, 1902
- Proagoderus gemmatus Peringuey, 1901 (BMNH drawer 218)
- Proagoderus gerstaeckeri Harold, 1867 (BMNH drawer 212)
- Proagoderus gibbiramus d'Orbigny, 1902 (BMNH drawer 212)
- Proagoderus hemicyaneus d'Orbigny, 1902 (BMNH drawer 218)
- Proagoderus ignitus d'Orbigny, 1905 (BMNH drawer 216)
- Proagoderus kilimanus Kolbe, 1891
- Proagoderus kolbei Marcus, 1921
- Proagoderus loricatus Klug, 1855 (BMNH drawer 217)
- Proagoderus lucasi d'Orbigny, 1902 (BMNH drawer 217)
- Proagoderus lujendae Bates, 1888 (BMNH drawer 216)
- Proagoderus mixtrifrons d'Orbigny, 1911 (BMNH drawer 217)
- Proagoderus nigricornis Fairmaire, 1887
- Proagoderus panoplus Bates, 1888 (BMNH drawer 215)
- Proagoderus prostans Reiche, 1847 (BMNH drawer 215)
- Proagoderus quadricristatus d'Orbigny, 1908
- Proagoderus ramosicornis d'Orbigny, 1902 (BMNH drawer 212)
- Proagoderus rangifer Klug, 1855 (BMNH drawer 212)
- Proagoderus rectefurcatus Fairmaire, 1891 (BMNH drawer 215)
- Proagoderus sexcornutus d'Orbigny, 1902 (BMNH drawer 215)
- Proagoderus speculicollis Quedenfeldt, 1884 (BMNH drawer 217)
- Proagoderus subextensus Kolbe, 1895 (BMNH drawer 216)
- Proagoderus tricornifrons d'Orbigny, 1908
PSEUDOCHIRONITIS
- Pseudochironitis stuhlmanni Kolbe, 1897 (BMNH drawer 172)
PSEUDOCOPRIS
- Pseudocopris tubericollis Felsche, 1910 (BMNH drawer 142)
PSEUDOSAPROECIUS
- Pseudosaproecius binasus d'Orbigny, 1907 (BMNH drawer 70)
- Pseudosaproecius mirepunctatus d'Orbigny, 1905
SAROPHORUS
- Sarophorus cicatricosus Peringuey, 1901 (BMNH drawer 73)
SCAPTOCNEMIS
- Scaptocnemis segregis Peringuey, 1901 (BMNH drawer 187)
SCARABAEOLUS
- Scarabaeolus ebenus Klug, 1855 (BMNH drawer 3)
SCARABAEUS
- Scarabaeus catenatus Gerstaecker, 1871 (BMNH drawer 8)
- Scarabaeus confusus Muller, 1942 (BMNH drawer 2)
- Scarabaeus fraterculus Kolbe, 1895 (BMNH drawer 10)
- Scarabaeus furcatus Castelnau, 1840 (BMNH drawer 10)
- Scarabaeus galenus Westwood, 1844 (BMNH drawer 8)
- Scarabaeus goryi Castelnau, 1840 (BMNH drawer 11)
- Scarabaeus interstitialis Boheman, 1857 (BMNH drawer 2)
- Scarabaeus isidis Castelnau, 1840 (BMNH drawer 13)
- Scarabaeus zambezianus Peringuey, 1901 (BMNH drawer 9)
SISYPHUS
- Sisyphus biarmatus Felsche, 1909 (BMNH drawer 41)
- Sisyphus costatus Thunberg, 1818 (BMNH drawer 40)
- Sisyphus crispatus Gory, 1833 (BMNH drawer 40)
- Sisyphus transvaalensis Peringuey, 1908 (BMNH drawer 40 & 41)
TANZANOLUS
- Tanzanolus leleupi Balthasar, 1960
TINIOCELLUS
- Tiniocellus spinipes Roth, 1851 (BMNH drawer 190)
TRICHONTHOPHAGUS
- Trichonthophagus fumatus d'Orbigny, 1913 (BMNH drawer 264)
- Trichonthophagus juvencus Klug, 1835 (BMNH drawer 259)

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